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An important effector of Arf6, Rab27a, and Rab35 is the GTPase activating protein (GAP) EPI64; the high concentration of EPI64 at the base of the microvilli binds to EBP50 of the ezrin/EBP50/F-actin complex (Reczek & Bretscher, 2001).
One study suggests that EPI64 lacks GAP activity for Arf6; overexpression of EPI64 augments levels of Arf-GTP (Hanono, Garbett, Reczek, Chambers & Bretscher, 2006).
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